Confirmation that spaces are really spaces

[Koen G]

Do spaces ever convey any information that could be required for interpreting the text, or are they wholly predictable?

My question would be: does the predictable writing system cause predictable spaces, or do spaces make the writing system more predictable? In the first case, spaces are spaces. In the second case, they may be something else.

it’s likely no coincidence that the glyph pairs with the highest incidences of unexpected behavior also tend to have relatively high proportions of ambiguous word breaks represented in the Zandbergen transcription, i.e., what I call “comma breaks,”

Might this be caused by transcribers feeling less certain about this situation than about, for example, an uncertain space after [n]?