Pareidolia, but underwater: What is under the green paint?

[Jorge_Stolfi]

Тry on this fragment:

Hm, unfortunately there is some bleedthrough of the dark brown ink of the other side (f22r) that may be responsible for those darker streaks on that clip:
A    
B    
C    
Image A is a clip of You are not allowed to view links. Register or Login to view. that covers yoru area of interest. Image B is a clip of the matching area on You are not allowed to view links. Register or Login to view. (as accurately as I could determine it), flipped left-to-right for convenient comparison.  Image C is image B with inverted colors and ~50% transparency overlaid on image A. 

On the left half of mage A, notice the faint ghosts of the berries of image B.  On image C, notice that there are many berries on the other side of your area of interest.

However, the ghosts are not that strong, so maybe they don't account for all the dark smudges in the area of interest.  I will see what I can do...

All the best, --stolfi