What's the evidence that the colors were added later?

[RobGea]

"given that there is now strong evidence that many of the bifolios and even quires were scrambled several times over the manuscript’s history and yet nearly all the paint transfers appear to be between pages in their current order, it seems that a great deal of the Voynich Manuscript’s paint was added later on in its life"

ciphermysteries --> "Voynich colour inference, a sure path to madness…"
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